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I suspect that in general, they are not, but perhaps someone who knows better could clarify.

However, I am looking specifically at the American Political Science Review, where contributions labeled "Letter" are much longer than is typical in other journals. Are these "Letters" peer reviewed?

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  • Ask them if you can't find it in their online materials. If the correct answer is "sometimes" then answers here might be misleading.
    – Buffy
    Jun 14, 2019 at 13:06

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Yes, they are peer-reviewed.

See also their author instructions:

The American Political Science Review (APSR) publishes letters, or research notes. [...] Although letters are reviewed like manuscripts [my own emphasis], they are shorter, more focused, demonstrate a novel perspective on existing research, and encourage scholarly debate [...].

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